Nut-cracking
| Insect-foraging | Algae-scooping
| Leaf-folding | Pestle-pounding
| Other
Bossou
chimpanzees on occasion use a probing stick to collect honey from the nest of sweat- or honey bees (Ohashi, 2006).
Bossou
chimpanzees customarily use a woody modified stick to prize adult
Xylocopa bees out of dead branches before reaching sometimes with a more slender probe for the larvae within.
Bossou
chimpanzees were observed to arrange large leaves on the ground as cushions
to sit on in order to avoid contact with the wet, moist ground beneath
(Hirata et al, 1998).
Chimpanzees at Bossou have been observed to insert sticks
or twigs into tree-holes or into the ground and then sniffing the tip
upon removal as a form of exploratory behavior.
While performing this behavior, a chimpanzee picks one
to several soft leaves, grasps the petiole between the thumb and the
index finger and then proceeds to repeatedly rip the leaf-blade with
its incisors, resulting in the production of a conspicuous and distinct
noise.
This behavior is clearly ritualized and serves as a communication signal.
The frequencies of mouth to finger leaf clipping vary across chimpanzee
communities and leaf clipping has not been observed in all communities
of chimpanzees studied thus far. For example, leaf clipping is absent
from the behavioral repertoire of the chimpanzees of Gombe. However,
it has been recorded at all other long-term study sites across Africa.
At Bossou and Mahale (K-group), exclusive use of the mouth for leaf
clipping has been recorded, whereas at other long-term field sites,
both techniques have been noted with varying frequencies (Whiten et
al, 1999).
This behavior has a communicatory function that is usually context-dependent.
Indeed, the signaling function of leaf clipping varies across communities.
At Bossou, the great majority of examples of leaf clipping is performed
to reflect clear frustration or in frustration-related aggression, while
only a few instances take place in a courtship context (Sugiyama, 1981).
During the habituation period of the Bossou chimpanzees, individuals
surprised in trees would leaf clip while watching the observer. Once
habituation had progressed, this form of leaf clipping considerably
deminished in frequency and is now mainly only seen in youngsters at
play.
| Aimed Throwing
during displays or other aggressive contexts |
Chimpanzees
at Bossou, as well as at many other study sites across Africa, have
been observed to throw a branch at another conspecific during display
or fighting contexts or at human observers who happened to come too
close!
Nut-cracking
| Insect-foraging | Algae-scooping
| Leaf-folding | Pestle-pounding
| Other